Deadline Info: Annually, Third Friday in January

Agency/Sponsor: National Science Foundation


The Division of Integrative Organismal Systems (IOS) supports research aimed at understanding why organisms are structured the way they are and function as they do. Proposals should focus on organisms as a fundamental unit of biological organization. Principal Investigators (PIs) are encouraged to apply systems approaches that will lead to conceptual and theoretical insights and predictions about emergent organismal properties. Areas of inquiry include, but are not limited to, developmental biology and the evolution of developmental processes, nervous system development, structure, and function, physiological processes, functional morphology, symbioses, interactions of organisms with biotic and abiotic environments, and animal behavior.


Preliminary Proposal (required): Third Friday in January, Annually  

Full Proposal: First Friday in August, Annually  

Award Information

Estimated Number of Awards: 220 

Anticipated Funding Amount: $55,000,000 

Eligibility Information

Proposals may only be submitted by the following: 

  • Universities and Colleges - Universities and two- and four-year colleges (including community colleges) accredited in, and having a campus located in, the US acting on behalf of their faculty members. Such organizations also are referred to as academic institutions.
  • Non-profit, non-academic organizations: Independent museums, observatories, research labs, professional societies and similar organizations in the U.S. associated with educational or research activities.
  • Consortia of only the eligible organizations listed above

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